Istiodactylus

Istiodactylus is a genus of a medium sized pterosaurs from the early Cretaceous period.

Description
Istiodactylus were moderately large pterosaurs. Correcting earlier estimates of a length of fifty-six centimetres, Mark Paul Witton in 2012 concluded that the skull of one specimen, NHMUK R3877, was forty-three centimetres long.[1] The maximum wingspan was probably about 4.3 metres (14.1 ft),[1] making the largest individuals roughly half the size of the largest Pteranodon. Because of the flat, rounded shape of the snout, Istiodactylus is sometimes referred to as a "duck-billed pterosaur".[2] However, unlike ducks, they had teeth. These triangular teeth were laterally compressed, an adaptation for slicing flesh, and interlocked tightly in the closed jaws. The skull was relatively short, with most teeth concentrated in the blunt front tip of the jaws. Witton's 2012 study proposed that Istiodactylus were primarily scavengers.

History and species
In 1887 Harry Govier Seeley named the remains of a fossil pelvis discovered on the Isle of Wight, Ornithodesmus cluniculus, thinking it was a bird.[3] In 1901 he considered that it might have been a pterosaur. In 1913 Reginald Walter Hooley named a second species, Ornithodesmus latidens, for some definitively pterosaurian material, found in the Vectis Formation. The holotype was BMNH R 0176, a partial skeleton. The specific name means "broad tooth" in Latin.

The type species of the genus Ornithodesmus, however, was in the 1980s discovered to be based on bones belonging to a dinosaur, which meant a new genus had to be named for O. latidens. This species was assigned to its own genus Istiodactylus by Stafford Howse, Andrew Milner, and David Martill in 2001. The genus name is derived from Greek istion, "sail" and daktylos, "finger", referring to the fact that the wing of pterosaurs is formed by a membrane attached to a wingfinger.

Howse et al. in 2001 created for Istiodactylus its own family Istiodactylidae.[5]

In 2006 a second species, I. sinensis from China, has been named, its specific name referring to China. Its holotype is NGMC 99-07-011, a partial skeleton of a subadult individual. It was much smaller than I. latidens, its dimensions being 63% of the larger species, thus about a quarter in weight.[6] In 2006 Lü Jun-Chang et al. concluded I. sinensis was a junior synonym of the istiodactylid Nurhachius.[7] Mark Witton has proposed it may be a synonym of Liaoxipterus.

Below is a cladogram showing the phylogenetic placement of this genus within Pteranodontia from Andres and Myers (2013).